Descriptions of koala fossils from the Miocene of Riversleigh, northwestern Queensland and implications for Litokoala (Marsupialia, Phascolarctidae)
JULIEN LOUYS1, KAREN BLACK1, MICHAEL ARCHER1,2, SUZANNE J. HAND1 AND HENK GODTHELP1
1School of Biological, Earth and Environmental Science, University of New South Wales, Sydney, New South Wales 2052 Australia
2Faculty of Science, University of New South Wales, Sydney, New South Wales 2052 Australia
Litokoala is revised on the basis of new material from the early to middle Miocene deposits of Riversleigh. A new species of Litokoala is described from the System B Outasite Local Fauna. Litokoala garyjohnstoni sp. nov. possesses a unique combination of both plesiomorphic and apomorphic dental features, making its phylogenetic position within Litokoala diﬃcult to interpret. In some aspects of dental morphology, L. garyjohnstoni is convergent on the early Miocene phascolarctid Nimiokoala greystanesi Black & Archer, 1997. The dentition of a partial skull of Litokoala from Jims Carousel Site, System C is also described. On the basis of this dentition, it is suggested that the morphological variation between L. kutjamarpensis Stirton, 1967 and L. kanunkaensis Springer, 1987 does not warrant speciﬁc distinction, the latter being the junior synonym.